The project Trajectories of Social-Ecological Systems in Latin American Watersheds: Facing Complexity and Vulnerability in the context of Climate Change (TRASSEwill identify which are the social (governance, economic, cultural) and biophysical dimensions whose interactions determine the trajectories in time of rural-urban watersheds SES in terms of measurable outcomes such as the flow of ecosystem services –in particular water regulation services-, land management strategies and human well-being. This will allow operationalizing the Social-Ecological System framework at the sub-watershed and watershed scales in order to predict what possible future trajectories might or might not be viable in terms of coping with climate change vulnerability and ensuring human well-being.